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SRX3465837: GSM2886483: C2C12D_T_2_3READS; Mus musculus; OTHER
1 ILLUMINA (NextSeq 500) run: 12.9M spots, 980.5M bases, 432.8Mb downloads

Submitted by: NCBI (GEO)
Study: Cellular stress alters 3'UTR landscape through alternative polyadenylation and isoform-specific degradation
show Abstracthide Abstract
Most eukaryotic genes express alternative polyadenylation (APA) isoforms with different lengths of 3' untranslated region (3'UTR). Here we show arsenic stress elicits global shortening of 3'UTRs through two mechanisms. First, stress leads to immediate shortening of 3'UTR due to preferential usage of proximal cleavage and polyadenylation sites (PASs), as revealed by 3' end sequencing of newly made RNAs that are metabolically labeled with 4-thiouridine. Second, long 3'UTR isoforms are more rapidly degraded during recovery from stress as compared to short 3'UTR isoforms, further shortening 3'UTR lengths in the cell. Using ribonucleoprotein immunoprecipitation coupled with 3' end sequencing (3'READS+RIP), we show that the RNA-binding protein T cell-restricted intracellular antigen-1 (Tia1) preferentially interacts with long 3'UTR isoforms via U-rich elements in alternative 3'UTR sequences, and the interaction correlates with stress granule (SG) association during stress and with mRNA decay during recovery from stress, indicating SG-mediated RNA clearance mechanism post stress. Importantly, genes whose 3'UTRs are shortened by APA during stress can evade stress-induced 3'UTR size-based mRNA degradation, leading to higher transcript abundance post stress. Moreover, proliferating and differentiated cells display different extents of 3'UTR shortening after stress, indicating cell type-specific of impact of stress on the 3'UTR landscape. Together, our data indicate that 3'UTR length plays important roles in gene expression in stressed cells, and APA functions as an adaptive stress response mechanism to preserve mRNAs. Overall design: 4 RNA-seq libraries from NIH3T3 cells for gene expression analysis; 4 3''READS libraries for analysis of APA using total RNA from NIH3T3 cells treated with sodium arsenite for 1h; 6 3''READS libraries for analysis of APA time course using total RNA from NIH3T3 cells treated with sodium arsenite for 1h and recovered for different amount of time; 8 3''READS libraries for analysis of APA isoform stability using 4sU metabolic labeling in NIH3T3 cells; 8 3''READS libraries for analysis of stress response of proliferating and differented C2C12 cells using cytoplasmic RNA; 4 3''READS libraries for the analysis of APA during C2C12 cell differentiation.
Sample: C2C12D_T_2_3READS
SAMN08166106 • SRS2753799 • All experiments • All runs
Organism: Mus musculus
Library:
Instrument: NextSeq 500
Strategy: OTHER
Source: TRANSCRIPTOMIC
Selection: other
Layout: SINGLE
Construction protocol: TRIzol 3'READS: The 3'READS procedure has been described in (Zheng et al., 2016). Briefly, Poly(A)+ RNA in 0.1-15 µg of total RNA was captured using 10 µl of oligo(dT)25 magnetic beads in 100 µl 1x binding buffer (10 mM Tris-Cl, pH7.5, 150 mM NaCl, 1 mM EDTA, and 0.05% TWEEN 20) and fragmented on the beads using 1.5 U of RNase III in 30 µl RNase III buffer (10 mM Tris-Cl pH8.3, 60 mM NaCl, 10 mM MgCl2, and 1 mM DTT) at 37 °C for 15 min. After washing away unbound RNA fragments with binding buffer, poly(A)+ fragments were eluted from the beads with TE buffer (10 mM Tris-Cl, 1 mM EDTA, pH 7.5) and precipitated with ethanol, followed by ligation to 3 pmol of heat-denatured 5' adapter (5'-CCUUGGCACCCGAGAAUUCCANNNN) in the presence of 1 mM ATP, 0.1 µl of SuperaseIn, and 0.25 µl of T4 RNA ligase 1 in a 5 µl reaction at 22 °C for 1 h. The ligation products were captured by 10 pmol of biotin-T15-(+TT)5 attached to 12 µl of Dynabeads MyOne Streptavidin C1. After washing with washing buffer (10 mM Tris-Cl pH7.5, 1 mM NaCl, 1 mM EDTA, and 0.05% TWEEN 20), RNA fragments on the beads were incubated with 0.01 U/µl of RNase H at 37 °C for 30 min in 30 µl of RNase H buffer (50 mM Tris-Cl pH 7.5, 5 mM NaCl, 10 mM MgCl2, and 10 mM DTT). After washing with RNase H buffer, RNA fragments were eluted from the beads in elution buffer (1 mM NaCl, 1 mM EDTA, and 0.05% TWEEN 20) at 50 °C, precipitated with ethanol, and then ligated to 3 pmol of heat-denatured 5' adenylated 3' adapter (5'-rApp/NNNGATCGTCGGACTGTAGAACTCTGAAC/3ddC) with 0.25 µl T4 RNA ligase 2 (truncated KQ version) at 22 °C for 1 h in a 5 µl reaction containing 15% PEG 8000 and 0.2 µl of SuperaseIn. The ligation products were then precipitated and reverse transcribed using M-MLV reverse transcriptase, followed by PCR amplification using Phusion high-fidelity DNA polymerase and bar-coded PCR primers for 13-18 cycles (13 cycles for 5 µg input, 15 cycles for 1 µg input, and 18 cycles for inputs below 1 µg). PCR products were size-selected twice with AMPure XP beads, using 0.6 volumes of beads (relative to the PCR reaction volume) to remove large DNA molecules and an additional 0.4 volumes of beads to remove small DNA molecules. The eluted DNA was selected again with 1 volume of AMPure XP beads to further remove small DNA molecules. The size and quantity of the libraries eluted from the AMPure beads were examined using a high sensitivity DNA kit on an Agilent Bioanalyzer. Libraries were sequenced on an Illumina NextSeq 500 (1x75 bases). RNA-seq: Poly(A)+ RNA in 5 µg of total RNA was purified twice using 10 µl of oligo(dT)25 magnetic beads (NEB) following the manufacturer's protocol. RNA was then fragmented by partial alkaline hydrolysis (94 °C for 2 min), 3' dephosphorylated with shrimp alkaline phosphatase, and 5' phosphorylated with T4 PNK. The RNA fragments were sequentially ligated to a 5' adenylated 3' blocked 3' adapter (5'-Aden/NNNNNNGATCGTCGGACTGTAGAACTCTGAAC/3ddC, where N is a random nucleotide) with RNA ligase 2 truncated, KQ (200 U/µl) and to a 5′ adaptor (5'-CCUUGGCACCCGAGAAUUCCA) with T4 RNA ligase I. Reverse transcription of ligation product, cDNA amplification by PCR, and library purification were performed as described above for the 3'READS procedure.
Experiment attributes:
GEO Accession: GSM2886483
Links:
Runs: 1 run, 12.9M spots, 980.5M bases, 432.8Mb
Run# of Spots# of BasesSizePublished
SRR637051012,901,771980.5M432.8Mb2018-06-13

ID:
4826193

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