| Modifier | Effect on White | Type of Regular |
|---|---|---|
| abnormal, small, homeotic discs 2 (ash2) | negative | transcription factor |
| apterous | negative | LIM family transcription factor |
| Beadex (Bd) | negative | transcription factor |
| brahma (brm) | positive | chromatin remodeling |
| cap-n-collar (cnc) | negative | bZIP transcription factor |
| devenir (dev) | negative | trithorax-Group |
| Distalless (Dll) | negative | transcription factor |
| Enhancer of Polycomb (E(Pc)) | negative | Polycomb Group |
| Enhancer of white-spotted[81d5] | positive | unknown |
| Enhancer of zeste (E(z)) | negative | Polycomb Group |
| extra sex combs (esc) | negative | Polycomb Group |
| hedgehog (hh) | positive | cell-cell signaling |
| Inverse regulator-a (Inr-a) | negative | unknown |
| kismet (kis) | positive | trithorax-Group |
| kohtalo (kto) | negative | trithorax-Group |
| l(2) 05208/Kruppel-homolog1 (Kr-h1) | positive | transcription factor |
| l(2) 03405/ Uba1 | positive | Ubiquitin activating |
| l(3) 00305 | negative | protein Ser/Thr kinase |
| l(3) 01969 | negative | unknown |
| l(3) 02104 | negative | unknown |
| l(3) 03670 | negative | alkaline phosphatase |
| l(3) 04063 | negative | unknown |
| l(3)04026 | negative | unknown |
| l(3)08232 (rolling pebbles) | negative | apoptosis inhibitor ? |
| l(3)87Ca/Vha55 | positive | Polycomb Group |
| Lightener of white (Low) | positive | unknown |
| mei-P19 | negative | cell cycle control? |
| Modifier of white (Mow) | negative | unknown |
| modulo (mod) | negative | DNA binding |
| osa (osa) | positive | DNA binding |
| oxen (ox) | negative | diacyl glycerol kinase |
| Polycomblike (Pcl) | negative | Polycomb Group |
| Posterior sex combs (Psc) | negative | Polycomb Group |
| Regena (Rga) | negative | transcription factor |
| Ribonuclear protein at 97D | negative | nuclear RNA binding |
| Ribosomal protein PO/AP lyase (PO) | negative | AP3 DNA endonuclease |
| scalloped (sd) | negative | transcription factor |
| Sex combs on midleg (Scm) | negative | transcription factor |
| skuld (skd) | negative | trithorax-Group |
| sugarless (sgl) | negative | UDP-G6DH (signaling) |
| Suppressor of Polycomb (Su(Pc)) | negative | Polycomb Group |
| Trithoraxlike (Trl) | positive | transcription factor |
| Ultrafemale overexpression (Ufo)/ lola | negative | transcription factor |
| urdur (urd) | negative | trithorax-Group |
| verthandi (vtd) | negative | trithorax-Group |
| Weakener of white (Wow) | positive/negative | unknown |
| Wingless (wg) | negative | cell-cell signaling |
From: Biological Consequences of Dosage Dependent Gene Regulation in Multicellular Eukaryotes
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