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Onthophagini
tribe, beetles
Nucleotide Protein
Scarabaeinae
(dung beetles & tumblebugs), subfamily, beetles
Copris
(black dung beetles), genus, beetles
Euoniticellus
genus, beetles
Onitis
Onthophagus
Digitonthophagus
Drepanocerus
Helictopleurus
Liatongus
Oniticellus
Phalops
Proagoderus
Tiniocellus
Tragiscus
Scarabaeinae incertae sedis
beetles
Oniticellini
Coprini
Microcopris
Serrophorus
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