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Onthophagini
tribe, beetles
Nucleotide Protein
Scarabaeinae
(dung beetles & tumblebugs), subfamily, beetles
Copris
(black dung beetles), genus, beetles
Euoniticellus
genus, beetles
Onitis
Onthophagus
Digitonthophagus
Drepanocerus
Helictopleurus
Liatongus
Oniticellus
Proagoderus
Tiniocellus
Tragiscus
Helictopleurus fulgens
species, beetles
Scarabaeinae incertae sedis
beetles
Oniticellini
Coprini
Microcopris
Serrophorus
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