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SRX2737273: GSM2577555: mock for CgPho2 ChIP w/ CgPho4 in high Pi media; [Candida] glabrata; ChIP-Seq
1 ILLUMINA (Illumina HiSeq 2000) run: 15.4M spots, 767.5M bases, 458.5Mb downloads

Submitted by: NCBI (GEO)
Study: Evolution of Reduced Co-Activator Dependence Led to Target Expansion of a Starvation Response Pathway [Cgla ChIP-seq]
show Abstracthide Abstract
In S. cerevisiae, the phosphate starvation (PHO) responsive transcription factors Pho4 and Pho2 are jointly required for induction of phosphate response genes and survival in phosphate starvation conditions. In the related human commensal and pathogen C. glabrata, Pho4 is required but Pho2 is dispensable for survival in phosphate-limited conditions and is only partially required for inducing the phosphate response genes. This reduced dependence on Pho2 evolved in C. glabrata and closely related species. Pho4 orthologs that are less dependent on Pho2 induce more genes when introduced into the S. cerevisiae background, and Pho4 in C. glabrata both binds to more sites and induces more genes with expanded functional roles compared to Pho4 in S. cerevisiae. We used Chromatin-ImmunoPrecipitation with exonucleas followed by high-throughput sequencing (BioChIP-seq) to identify the binding locations of Pho4 from both S. cerevisiae and C. glabrata in the S. cerevisiae background lacking the negative regulator Pho80, and either with or without Pho2. Overall design: We made C. glabrata strains carrying a C-terminally tagged (3x FLAG) version of the endogenous Pho4 either with or without the endogenous Pho2. We also made strains carrying an N-terminally tagged (V5) version of the endogenous Pho2. Mock strains were made to correspond to the experimental strain's genetic background, but lacking the epitope tag. All strains were grown to mid-log phase, washed and released into no phosphate media for 1 hour and harvested for ChIP analysis. We used ChIP-exo, a variation of the ChIP protocol using exonuclease to increase the resolution and signal-to-noise ratio, to query the genomic binding locations of both Pho4 and Pho2 in C. glabrata. A routinely used control for ChIP is the input chromatin for normalizing against heterogeneity in the fragmentation process. This, however, doesn't apply to the ChIP-exo technique. We use either hte mock sample instead for peak finding and generating the WIG files.
Sample: mock for CgPho2 ChIP w/ CgPho4 in high Pi media
SAMN06756868 • SRS2124574 • All experiments • All runs
Library:
Instrument: Illumina HiSeq 2000
Strategy: ChIP-Seq
Source: GENOMIC
Selection: ChIP
Layout: SINGLE
Construction protocol: The cross-linked cell sample was resuspended in ~1 mL of ChIP lysis buffer (50 mM HEPES, pH 7.5, 140 mM NaCl, 1 mM EDTA, 1% Triton X-100, 0.1% Na-Deoxycholate), and mechanically lysed using a mini-Beadbeater-24 (BioSpec) with 0.5 mm Zirconia beads (BioSpec), using the following setting: 12 cycles of 30s bead beating, with 1 min on ice-water bath in between. The lysed sample was fragmented by sonication using Covaris E220 Adaptive Focus system (setting: Duty Factor: 10%; Peak Incident Power: 175; Cycles/Burst: 200; Time:150s). Afterwards, the sample was centrifuged to remove the unsoluable fraction. The supernatant containing the solubalized chromatin was subjected to immunoprecipitation. Briefly, the supernatant was incubated with either Anti-FLAG M2 magnetic beads (Sigma-Aldrich) or Anti-V5 antibody (Abcam #15828) depending on the epitope tag that was used. After 2 hours of incubation at 4C with rotation, the V5-tagged samples were additionally incubated with Dynabeads Protein-G (Thermo-Fisher Scientific #10003D). In a traditional ChIP experiment, what follows are the washing and de-crosslinking, prior to DNA extraction. In ChIP-exo, however, the washing steps are combined with part of the library construction steps. A detailed account for the protocol can be found in (Rhee and Pugh, 2012), and is also available from Active Motif ChIP-exo vA4 manual. As mentioned above, the ChIP washing step is combined with part of the library construction steps in ChIP-exo. A brief description of the steps is provided here: with the chromatin still bound to the beads, the DNA was end-polished and P7-exo adapters are ligated onto the blunt ends. The nicked DNA was repaired and then digested by lambda and RecJf exonulceases to excise DNA in a 5’ to 3’ direction, trimming up to the site of the cross-linking and selectively eliminating the P7 adapter at the 5 ́ end. Following cross-link reversal and elution from the beads, the DNA was made double-stranded by P7 primer extension and a P5- exo adapter was added to the exonuclease-treated ends. The DNA library was PCR amplified and size selected before it was subjected to high-throughput sequencing.
Experiment attributes:
GEO Accession: GSM2577555
Links:
Runs: 1 run, 15.4M spots, 767.5M bases, 458.5Mb
Run# of Spots# of BasesSizePublished
SRR544840615,350,616767.5M458.5Mb2017-04-18

ID:
3945731

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