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SRX18348507: GSM6751466: JW1-SR-H3K27me3; Triticum aestivum; ChIP-Seq
1 ILLUMINA (Illumina NovaSeq 6000) run: 38.1M spots, 11.4G bases, 3.5Gb downloads

External Id: GSM6751466_r1
Submitted by: Functional Epigenomics Group, National Key Laboratory of Plant Molecular Genetics, Institute of Plant Physiology and Ecology,Chinese Academy of Sciences
Study: LHP1-mediated conditional repression of subgenome-diversified defenses is responsible for the high plasticity of allopolyploid wheat
show Abstracthide Abstract
High plasticity of common wheat is attributed to the captured and polyploidization-promoted diversity. However, uncontrolled subgenome diversification can lead to hybrid conflict and dysgenesis, resulting in decreased diversity. How genomic diversity is maintained and interpreted to increase plasticity is unclear. By data-mining from the binding of 193 genome-wide trans-factors and genetic perturbations in common wheat, we identified LHP1 as a major regulator of subgenome-diversified defense genes, enhancer RNAs, and metabolite synthesis-related gene clusters via H3K27me3. Stripe rust infection leads to a global decrease in LHP1-mediated H3K27me3, deprivation of which enhances common wheat stripe rust resistance. We also revealed the consistency between subgenome diversity and population diversity, potentially promoted by LHP1, implying the recent diversification preferentially occurred in the captured subgenome-diversified regions regulated by LHP1. Thus, common wheat benefitted from multi-faced role of LHP1 in promoting sequence diversity and repressing subgenome-diversified defenses; this constraint is eliminated by pathogen infections, enabling timely release and fixation of favorable variations, conferring the evolutionary advantage and high plasticity of common wheat. Overall design: To elucidate how genomic diversity is maintained and interpreted to increase plasticity in common wheat, we obtained gene expression profiles, H3K27me3, LHP1 and eRNA in wild type (JW1) and lhp1-abd mutant.
Sample: JW1-SR-H3K27me3
SAMN31836741 • SRS15836819 • All experiments • All runs
Library:
Name: GSM6751466
Instrument: Illumina NovaSeq 6000
Strategy: ChIP-Seq
Source: GENOMIC
Selection: ChIP
Layout: PAIRED
Construction protocol: Chromatin or RNA was extracted from 14-days-old seedlingss, using standard protocols. For RNA-seq, library construction and deep sequencing were performed by Novogene (Beijing, China) using Illumina NovaSeq 6000 system to produce 150-bp paired-end reads. For ChIP-seq, crosslinked materials were ground into fine power with liquid nitrogen and resuspended in ChIP Lysis Buffer1 (CLB1: 50 mM HEPES [pH 7.5], 150 mM NaCl, 1 mM EDTA, 1% Triton X-100, 10% Glycerol, 1x inhibitor Cocktail, 0.035% 2-Mercaptoethanol) and incubated for 60 minutes with rotation at 4°C. After incubation, collect the nucleus after filtering the mixture through 40-um strainer, centrifuging at 3000 g for 30 minutes at 4°C in a swinging bucket rotor and removing the supernatant. The nucleus was washed twice with ChIP Lysis Buffer2 (CLB2: 50 mM HEPES [pH 7.5], 150 mM NaCl, 1 mM EDTA, 1% Triton X-100, 10% Glycerol, 1x inhibitor Cocktail). DNA was sheared by sonication to approximately 300–500-bp fragments. After centrifugation (10 minutes at 13,000 rpm), the supernatant was precleared with 40 μl salmon sperm (SS) DNA/Protein A agarose for 60 minutes at 4°C. After 2 minutes of centrifugation at 500 g, the supernatant was transferred to a siliconized tube, and 10 μl of the appropriate antibody (H3 trimethyl-Lys 27 (ABclonal, A2363), H3 trimethyl-Lys 4 (Abcam, Cambridge, England), and H3 acetyl-Lys 9 (Millipore)) was added. After incubation overnight with rotation, 40 μl SS DNA/Protein A agarose was added and incubation continued for 1 hr. The agarose beads were then washed with 1 ml of each of the following: Low salt buffer (50 mM HEPES [pH 7.5], 150 mM NaCl, 1 mM EDTA), High salt buffer (50 mM HEPES [pH 7.5], 635 mM NaCl, 1 mM EDTA), LiCl wash buffer (0.25M LiCl, 0.5% NP-40, Tris-HCl [pH 8], 1 mM EDTA), and 1× TE (10 mM Tris-HCl [pH 8], 1 mM EDTA). The immunocomplexes were eluted from the beads with 400 μl 1% SDS, 0.1 M NaHCO3. A total of 20 μl 5 M NaCl was then added to each tube, and crosslinks were reversed by incubation at 65°C for 5–6 hours. Residual protein was degraded by the addition of 20 μg Prot K (in 10 mM EDTA and 40 mM Tris [pH 8.0]) at 45°C for 1 hour, followed by phenol/chloroform/isoamyl alcohol extraction and ethanol precipitation. Pellets were washed with 70% EtOH and resuspended in 30 μl TE buffer. More than 10 ng ChIP DNA was used to prepare each sequencing sample. Libraries were constructed and sequenced by Novogene (Beijing, China). The libraries were sequenced with the Illumina NovaSeq 6000 system to produce 150-bp paired-end reads. For spike-in ChIP-seq, Arabidopsis chromatin was added into wheat chromatin before immunoprecipitation. Briefly, before performing immunoprecipitation, 40 ul of chromatin was reverse crosslinked, proteinase K digested followed by DNA purification. The concentration of purified DNA was determined using Qubit dsDNA High-Sensitivity Assays (Invitrogen). Meanwhile, chromatins from Arabidopsis were also isolated and determined the concentration with the same method. One percent of Arabidopsis chromatins were mixed with different Wheat samples before immunoprecipitation. The rest procedures are the same as the previous description. For nascent RNA (chromatin-bound RNA, CB RNA) library preparation, the chromatin RNA extraction was completed as previously described54. After degrading genomic DNA by TURBO DNase (Life Technologies), CB RNA was subjected to rRNA depletion using a riboPOOL kit (siTOOLs Biotech, PanPlant-10 nmol) and polyA RNA removal by oligo(dT) beads (NEB, S1419). Both CB RNA and polyA RNA were transformed into cDNA libraries using the NEBNext Ultra II Directional RNA Library Prep Kit for Illumina (NEB #E7765) and sequenced on an Illumina NovaSeq platform.
Runs: 1 run, 38.1M spots, 11.4G bases, 3.5Gb
Run# of Spots# of BasesSizePublished
SRR2237798238,089,99311.4G3.5Gb2023-10-12

ID:
25362726

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