1 Department of Cell Biology, Yale School of Medicine, New Haven, CT 06520, United States. Electronic address: yongli.zhang@yale.edu.
2 Department of Biophysics and Biophysical Chemistry, Johns Hopkins University, Howard Hughes Medical Institute, Baltimore, MD 21205, United States; Department of Biophysics, Johns Hopkins University, Howard Hughes Medical Institute, Baltimore, MD 21205, United States; Department of Biomedical Engineering, Johns Hopkins University, Howard Hughes Medical Institute, Baltimore, MD 21205, United States. Electronic address: tjha@jhu.edu.
3 Department of Molecular & Cell Biology, Institute for Quantitative Biosciences (QB3)-Berkeley, University of California, Berkeley, Berkeley, CA 94720, United States. Electronic address: marqusee@berkeley.edu.