1 Department of Earth, Ocean and Ecological Sciences, University of Liverpool, Liverpool, L69 3GP, UK; Department of Zoology and Entomology, University of Pretoria, Pretoria 0002, South Africa. Electronic address: T.R.Bishop@liverpool.ac.uk.
2 Department of Earth, Ocean and Ecological Sciences, University of Liverpool, Liverpool, L69 3GP, UK. Electronic address: Hannah.griffiths@liverpool.ac.uk.
3 School of Biological Sciences, The University of Hong Kong, Hong Kong, SAR, China.
4 Department of Life Sciences, Natural History Museum, London, UK.
5 Department of Earth, Ocean and Ecological Sciences, University of Liverpool, Liverpool, L69 3GP, UK; Department of Life Sciences, Natural History Museum, London, UK.
6 Department of Earth, Ocean and Ecological Sciences, University of Liverpool, Liverpool, L69 3GP, UK; Department of Zoology and Entomology, University of Pretoria, Pretoria 0002, South Africa; School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Wits, South Africa.